Alias used by several supervillains appearing in American comic books published by Marvel Comics, usually depicted as enemies of the superhero Spider-Man. The original and best known incarnation, Miles Warren, was originally introduced in The Amazing Spider-Man #31 as a professor at the fictional Empire State University. Later storylines established him as also being a scientist researching genetics and biochemistry, and revealed an unhealthy romantic obsession he had for Gwen Stacy. Warren was driven mad with grief and jealousy so he created his Jackal alter-ego to seek revenge on Spider-Man, whom he blamed for Gwen's tragic death..